the pen shell (pinna nobilis) ................................................................................................................................................................................................

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the pen shell (pinna nobilis)
- august 2002 -




the swedish naturalist carl linnaeus, also known as
carl von linné or carolus linnaeus (1707-1778)
first described the species in 1758.
it is an endemic species of the mediterranean,
belonging to the family of pinnidae.
this large, brown fan-shaped bivalve shellfish,
one of the largest in the world,
can reach a breadth of 1,2 metres.
its habitat is in the coastal region, on the shoals
that are either sandy or overgrown by sea-flowering plants.
it is found mainly in south italy /sardinia, but also in
france /around the mediterranean islands (corsica ...),
on the coasts of yugoslavia, in north-africa, etc.

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byssus threads
the pen shell has a byssus gland that secretes byssus,
an organic maner that hardens in contact with water
and forms threads that help fasten the valves
to the base. the fan mussel lives half buried in
the sediment where it finds nourishment and shelter.
like many other organisms it remains anached to the
same substratum in the same place, almost stationary,
to the end of its life.
it can, however, use its foot to move slowly and in
limited fashion. this means that the fan mussel can be
found at different depths depending on age.
during the larva stadium it spends its life swimming freely.
the young individuals being found near the surface,
while the older individuals live at a depth of approximately
40 metres.

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the decline in populations
threatened with extinction for some forty years -
it is partly due to the decline in the posidonia oceanica
seagrass meadows.
trawling, anchoring of yachts, various pollutants, sea resort
and other developments, as well as unauthorized fishing by
certain divers, have all contributed to accelerating the
decline of this species.
the decline in the seagrass has serious implications for
the fan mussel whose highly specific ecology and
metabolism renders its reproduction difficult.

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reproduction
the eggs mature and the larvae develop in the open seas.
then, made heavy by a thin calcitic shell following its planktonic
phase, the larva falls to the bottom where it attaches itself.
at this stage it measures approximately 2 cm and has a
transparent and fragile shell, which leaves the animal prey
to numerous predators, such as octopus, sea bream, etc.
the mortality rate at this stage is therefore high.


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not only were molluscs used to dye cloth,
they were also used to manufacture cloth.
ancient artisans used the fine strong bysus threads
of the noble pen shell to manufacture the legendary
sea silk.
see the history of byssus cloth


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chiara vigo in sant'antioco, sardinia
is one of the few women today, who are
still manufacturing byssus cloth.
see more


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it must be spinned according to the mooon
see how to manufacture byssus threads





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the pen shell (pinna nobilis) with
tuft of silky byssus threads




fishing of the pinna nobilis




the pen shell is dug into the oozy-sandy base
with a third of its valve by its sharp pointed end




pinna nobilis, linné, 1758